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The other save I have is the same way. Mountain biome doesn't allow for saplings; Temperate Freshwater Marsh does. Now that I look at the list again I think the mountain biome is actually blocking a lot seeds. Only 10 seed types can be planted in the Mountains vs like 30+ otherwise.
I don't think either get freezing temp (both have a river that does not freeze in winter). Do mountains just count as a freezing biome by default maybe?
So between my two saves I can only plant trees on fire clay.